Robert Warren Howarth

Department of Ecology & Evolutionary Biology
Cornell University, USA
E-mail: rwh2(at)

Fields of interest:

  • Biogeochemistry and aquatic ecosystem science;
  • global and regional nitrogen and phosphorus cycles; 
  • biotic, physical, and geochemical controls on nitrogen fixation; 
  • influence of land-use, management practices, and climate change on nutrient fluxes from the landscape; 
  • atmospheric deposition of nitrogen onto the landscape; 
  • controls and consequences of eutrophication in estuaries; 
  • environmental management and the effects of pollutants on aquatic
  • environmental consequences of biofuels; 
  • application of science to sustaining the biosphere.


Recent publications:

R.W Howarth, H. Paerl: 
Coastal marine eutrophication: Control of both nitrogen and phosphorus is necessary. 
Proceedings of the National Academy of Sciences, Vol. 105, 2008, 
p. 103

R.W. Howarth: 
The nitrogen cycle.
In: G. E. Likens (ed.), Encyclopedia of Inland Waters. Elsevier, Oxford, 2009

R. Marino, R.W. Howarth: 
Nitrogen fixation.
In: G. E. Likens (ed.), Encyclopedia of Inland Waters. Elsevier, Oxford, 2009

R.W. Howarth: 
Coastal nitrogen pollution: A review of sources and trends globally and regionally. 
Harmful Algae Vol. 8, pp. 14-20

E.W. Boyer and R. W. Howarth: 
Nitrogen fluxes from rivers to the coastal oceans. 
In: D. Capone, D. A. Bronk, M. R. Mulholland & E. J. Carpenter (eds.), Nitrogen in the Marine
, 2nd Edition, Elsevier, Oxford, 2008

D.P. Swaney, D. Scavia, R.W. Howarth, R. M. Marino: 
Estuarine classification and response to nitrogen loading: Insights from simple ecological models. 
Estuarine,Coastal & Shelf Science Vol. 77, pp. 253-263

T.W. Simpson, A.N. Sharpley, R.W. Howarth, H. W. Paerl, K.R. Mankin: 
The new gold rush: Fueling ethanol production while protecting water quality. 
Journal of Environmental Quality Vol. 37, 2008, pp. 318-324

R. Marino, F. Chan, R.W. Howarth, M.L. Pace, G.E. Likens: 
Ecological constraints on planktonic nitrogen fixation in saline estuaries. I. Nutrient and trophic controls, 
Marine Ecology Progress Series Vol. 309, 2006, pp. 25-39

F. Chan, M.L. Pace, R.W. Howarth, M. Roxanne:
Bloom formation in heterocystic nitrogen-fixing cyanobacteria: The dependence on colony size and zooplankton grazing, 
Limnology and Oceanography Vol. 49(6), (2004), pp. 2171-2178